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YKTTW should be done as a forum.
YKTTW, as it's currently set up, is best if you have a fully-formed idea for a trope that just needs examples and minor touching-up and the approval of several other tropers to get launched. It's not very good if you just have a vaguely-formed idea and want to get comment to flesh it out.
The YKTTW system is very focused on showing the topline or opening post stuff. The proposal is the biggest thing. If people are skimming, they'll first see the latest comments (unless they haven't read them at all), and then the proposal (with the first few lines being the most likely visible), and then old comments.
The main YKTTW page--as much as it wants people to contribute to other people's ideas--does not do a good job at encouraging wading into others YKTTWs. Each YKTTW takes up a lot of space on the screen--you could make the page more easily skimmable if you had more on screen at once, such as in a typical forum format where at most you have two small lines for each YKTTW, a title and a laconic description.
Now, if you say that people ought to always pay attention to updates in the trope description when they're going to look at new comments, I can say that you can specially code the YKTTW forum so that people aren't directed to the last post they've read, but always to the top. People can still hit End on their keyboard, but they will at least have a chance to catch any special bolded text yelling at them in the brief moment after the page first loads.
Comments
I think you bring up a good point, but from what I hear, there are many more people who just use the TV Tropes wiki than the number who even use the forum at all, so making YKTTW a forum may mean fewer people contributing to it. That being said, it is probably a good idea to take what I say with a grain of salt here since I do not have extensive experience with YKTTW.
Still, I definitely agree about YKTTW being better for proposals that are near completion than for vague ideas about tropes. I am not sure that it is currently set up to encourage much in the way of substantive cooperation and I think that is kind of a problem in TRS and Special Efforts too.